Origins
Starting conditions and restrictions.
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61 origins
Arc WeldersHomeworld: Initial 
Mining Districts replaced with Generator DistrictsThe Machine Age
Broken RuleSpecies Traits: Perfected Genes Ruler Traits: Luminary
Broken ShacklesInitial 
Pops: Species from Pre-FTL Civilizations across the galaxyFirst Contact Story Pack
Calamitous BirthHomeworld: Massive Crater 
Buried Lithoids: +4Lithoids Species Pack
Clone ArmyHomeworld: Ancient Clone Vat: +2 Species Traits: Clone Soldier
Common GroundLeader of a non-Hegemony Federation Guaranteed Habitable Worlds: Replaced with Federation membersFederations
Cosmic DawnHomeworld: Volcanic World (Size: 14) Ancient Stasis Pods: +5Infernals Species Pack
Cybernetic CreedHomeworld: Initial buildings replaced by Creed InfrastructureThe Machine Age
DoomsdayWARNING: Challenging Origin This civilization's homeworld is highly unstable, and it is only a matter of time before it explodes. Their only hope is to seek refuge elsewhere before it is too late.Federations
EndbringersWARNING: Challenging Origin WARNING: Blocks all Ascension Paths except for Psionic Ascension.Shadows of the Shroud
Evolutionary PredatorsSpecies Traits: Malleable Genes Situation: Adaptive EvolutionBioGenesis
Fear of the DarkHome System: Pre-FTL Civilization created by a Separatist Faction, formerly part of your Empire, granting events and benefitsFirst Contact Story Pack
Forever CruiseNomadic Origin Start with Situation: Passenger SatisfactionNomads
Fruitful PartnershipSpace Fauna: Disperse Seed Pods throughout the galaxyPlantoids Species Pack
Galactic DoorstepHome System: Dormant Gateway
Hard ResetHomeworld: Colony Modifiers: Battlefield Recoveries
HegemonLeader of a Hegemony Federation Guaranteed Habitable Worlds: Replaced with Federation membersFederations
Heirs of the KhanNomadic Origin Periodically chased by Hostile Marauder shipsNomads
Here Be DragonsHome System: Inhabited by a protective Sky Dragon, that will become wrathful if displeasedAquatics Species Pack
Imperial FiefdomSubject of an Advanced AI Empire Can select a Specialist Subject type at the start of the gameOverlord
Knights of the Toxic GodHomeworld: Toxic Planetary Features Unique Orbital Habitat:Toxoids Species Pack
Life-SeededHomeworld: Gaia World (Size: 30) with Rare Planetary FeaturesApocalypse
Lost ColonyStarting Planet: Colonial Spirit Planetary Modifier
MechanistRobot Pops: 800 Mechanical Trait: Adaptive Frames Homeworld: 
Robot Assembly Plant
MindwardensAdditional Starting Technologies: 
Starhold Home System: Start with a Starhold.Shadows of the Shroud
NecrophageHomeworld: Chamber of Elevation Species Traits: NecrophageNecroids Species Pack
Ocean ParadiseSpecies Traits: Aquatic Homeworld: Ocean World (size: 30)Aquatics Species Pack
On the Shoulders of GiantsHome System: Ex Gravitas Archaeology Site related to a mysterious benefactorFederations
OvertunedSpecies Traits: Access to Overtuned Traits Overtuned Traits: Can be boosted by the Damn the Consequences EdictToxoids Species Pack
PaybackHomeworld: Advanced Infrastructure Colony Modifiers: Unified PurposeFirst Contact Story Pack
Post-ApocalypticHomeworld: Tomb World Species Traits: SurvivorApocalypse
Primal CallingHomeworld: Colony Modifiers: Untamed Wilds Available Buildings: Wildlife RanchGrand Archive
Progenitor Hive Spawning Pools replaced with Offspring Nest Available Holdings: Offspring NestOverlord
Prosperous UnificationInitial Pops: +400 Homeworld: Generator Districts: +1
Radioactive RoversHomeworld: Tomb World Species Traits: Radiation ShieldsApocalypse
Red GiantHomeworld: Volcanic World (Size: 20)Infernals Species Pack
RemnantsHomeworld: Relic World (Size: 22) Ruined Arcology: +6 …Ancient Relics Story PackFederations
Resource ConsolidationHomeworld: Prototype Machine World Planetary Features: Consolidated Resources
RiftworldHomeworld: Ancient Crater Archaeology Site Colonized Planets in systems with Astral Rifts: Rift AffinityAstral Planes
Scion
Scion of a Fallen Empire Overlord Home System: Connected to Overlord's capital via a WormholeFederations
Shattered RingHomeworld: Shattered Ring Segment Guaranteed Habitable Worlds: Replaced with Colonizable Ring World Segments that have Blockers decreasing HabitabilityFederations
Shroud-ForgedHomeworld: Shroud-Touched Region: +4 Start with 1000 
Shroud-Forged PopsShadows of the Shroud
Slingshot to the StarsHome System: Nearby ruined Quantum Catapult Twin Arrays with the Logs from the Quantum Architect Archaeology SiteOverlord
Starlit CitadelHome System: Deep Space Citadel I at the edge of the gravity wellBioGenesis
Storm ChasersLeader Traits: Storm Rider Planet 
Unity from 
Jobs per Storm Building: +5%Cosmic Storms
Subaquatic MachinesSpecies Traits: Waterproof Homeworld: Ocean World (size: 30)Aquatics Species Pack
SubterraneanSpecies Traits: Cave Dweller Mining Districts: UncappedOverlord
Subterranean MachinesSpecies Traits: Molebots Mining Districts: UncappedOverlord
Syncretic Evolution Organic Pops with Serviles Trait: 2.4K
Synthetic FertilityInitial Pops: +900 Species Traits: Pathogenic GenesThe Machine Age
Teachers of the ShroudSpecies Traits: Latent Psionic Available Starbase Buildings: Shroud BeaconOverlord
The Sacred PathNomadic Origin Unique Arkship District: Consecrated CommunityNomads
Treasure HuntersStarts with an inactive Celestial Chart Every recruited leader has a bonus Adventurous Spirit traitGrand Archive
Tree of LifeHomeworld: Tree of Life Colonized Planets: Tree of Life Sapling Planetary Feature
Under One RuleSpecies Traits: Perfected Genes Ruler Traits: LuminaryGalactic Paragons
Under One RuleSpecies Traits: Perfected Genes Ruler Traits: Luminary
Under One RuleSpecies Traits: Perfected Genes Ruler Traits: Luminary
Void DwellersHomeworld: Orbital Habitat Species Traits: Void DwellerFederations
VoidfarersNomadic Origin Start with an additional Logistic ShipNomads
VoidforgedHomeworld: Orbital Habitat Species Traits: Zero-G OptimizedFederations
WildernessSpecies Trait: Wilderness Replaces all standard Buildings, Districts, and 
District Specialization with bespoke onesBioGenesis